8  First signs of a zodiacal indicator marking up the horizontal angle (7)

 9  Biblical character is a retired accountant (5)

10 South America's fencing in cavity (5)

11 Copy one couple without it (7)

12 Unexpectedly thrives without five of them (6)

14 One deadly sin is acceptable (6)

16 Refined gelatine mixture without pig guts (7)

17 Vegetable is reportedly extremely overcooked (5)

20 Excuse retrograde sibilance to some extent (5)

21 Fifty-six breaking into study and starting to go rummaging about (7)

22 Inclusion commas sent pen crazy (13)

 1  Waterproof, primarily subaquatic creatures (5)

 2  Particle accelerator now interests our nonchalant leaders (5)

 3  Riding it means risque activity (13)

 4  Chetnik scratching bottom? Frightfully cultural (6)

 5  Varying measures of class I singled out (7,6)

 6  Oriental guy munched on stem (7)

 7  Come through draw against top player, by the sound of it (7)

12 Generate confusion after Romeo flees juvenile (7)

13 Sorrowful eastern Gaelic composition (7)

15 Way one of Verdi's operas lifts arenas (6)

18 Female relative discussed going after original aromatic spice (5)

19 Figure it must come after love (5)